Justine Greening, who is opposed to a third runway at Heathrow, has been replaced as transport secretary by Patrick McLoughlin.
"There can be only one reason to move her - and that is to expand Heathrow Airport," Mr Johnson said.
The idea was "mad" and he would fight it all the way, he added.
In a statement, the mayor said: "The third runway would mean more traffic, more noise, more pollution - and a serious reduction in the quality of life for hundreds of thousands of people.
"We will fight this all the way. Even if a third runway was built, it would not do the job of meeting Britain's needs.
"If we are to remain Europe's premier business hub we need a new four-runway airport, preferably to the east of London, that addresses the problem of aviation capacity before it is too late, and business is driven into the arms of our European competitors.
"It is time for Patrick McLoughlin to look at all the options, including bolder solutions that would deliver massive benefits in jobs and growth.
"And it is time for the government to level with Londoners. Are they in favour of a third runway at Heathrow or not?"
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